cAMP regulates IL-10 production by normal human T lymphocytes at multiple levels: A potential role for MEF2

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Συγγραφείς

Liopeta, K.
Boubali, S.
Virgilio, L.
Thyphronitis, G.
Mavrothalassitis, G.
Dimitracopoulos, G.
Paliogianni, F.

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peer reviewed

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Mol Immunol

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Περιγραφή

Signal transduction by the cAMP/cAMP-dependent protein kinase A (PKA) pathway is triggered through multiple receptors and is important for many processes in a variety of cells. In T cells, the engagement of the TCR-CD3 complex induces CAMP, a second messenger that controls immune response. IL-10, produced by a variety of lymphocyte subpopulations, is an important regulator of this response exerting a wide range of immunomodulatory actions. Elevation of cAMP has been shown to increase IL-10 production by monocytes. However, the mechanism of cAMP mediated regulation of IL-10 production by T lymphocytes remains unclear. In this study using normal peripheral T lymphocytes stimulated either through the TCR-CD3 complex or the TCR-CD3 and the CD28 molecule, we show that IL-10 is produced mainly by memory T lymphocytes after either way of stimulation and is drastically inhibited (70-90%) by CAMP elevating agents. cAMP mediated inhibition was reversed by the use of the specific PKA inhibitor Rp-8-Br-cAMP but not by the addition of exogenous rhIL-2, indicating that the inhibitory effect depends on PKA activation and is not secondary to IL-2 inhibition, Inhibition is taking place at both transcriptional and posttranscriptional level. Transfection of a luciferase reporter plasmid carrying the IL-10 promoter in T cells, revealed that TCR/CD28-induced activation was inhibited by 60% by cAMP elevation. The most sensitive part to cAMP mediated inhibition was a fragment of 135 bp Upstream of TATA box, which contains multiple binding sites for MEF-2. Overexpression of MEF-2 in the same cells increased IL-10 promoter activity by 2.5-fold. Stimulation through TCR/CD28 increased MEF-2 binding in its Corresponding binding sites which was inhibited by 80% in the presence of cAMP elevating agents. These results suggest that the inhibitory effect of cAMP on IL-10 production by normal peripheral T lymphocytes is cell type and stimulus specific, exerted on multiple levels and involves MEF2 transcription factor. (C) 2008 Elsevier Ltd. All rights reserved.

Περιγραφή

Λέξεις-κλειδιά

camp, il-10, t lymphocytes, mef2, pka, myocyte-enhancer factor-2, protein-kinase-a, necrosis-factor-alpha, gene-expression, transcription factors, interleukin-10 promoter, cell-proliferation, muscle development, prostaglandin e-2, binding protein

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<Go to ISI>://000262810400005
http://ac.els-cdn.com/S0161589008007438/1-s2.0-S0161589008007438-main.pdf?_tid=44e2cca8468d07c16b2bdd726b6f207a&acdnat=1336730790_1fe974404445ea4d356893933d908dab

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en

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Γενική Περιγραφή / Σχόλια

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Πανεπιστήμιο Ιωαννίνων. Σχολή Επιστημών και Τεχνολογιών. Τμήμα Βιολογικών Εφαρμογών και Τεχνολογιών

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